Monograph of Palaearctic, Oriental, and New Guinean Pinophilina (Coleoptera: Staphylinidae: Paederinae)

Assing, V.


Supplement of Volume 107, 2022
Published on 15 July 2022
ISBN 978-80-7028-570-1
ISSN 1211-8788

Contact:

dr. Jiří Kolibáč (Head of the Department of Entomology), email: jkolibac@mzm.cz
Milan Mačinec (Head of the Publishing Department), email: mmacinec@mzm.cz

Currently including more than 380,000 species, beetles are by far the most speciose of all organism orders and represent nearly one fourth of our planet´s known biodiversity. With a total of ~66,000 described extant species, rove beetles (Staphylinidae) are probably the most biodiverse family not only of beetles, but of all organisms on Earth, and hundreds of new species are added every year. Staphylinidae are distributed worldwide and ecologically remarkably diverse. They are found in practically all terrestrial habitats from wet to arid, from sea-level to high-alpine, and from deep soil to the canopy layer of forests. Numerous species are inquilines living with ants, termites, wasps, mammals, and birds or inhabitants of mushrooms and various kinds of decaying matter such as compost, dead wood, carrion, and animal excrements. Despite the important role of rove beetles in practically all terrestrial ecosystems, relatively little is known about their life histories and ecology. Read more...
  • Monograph of Palaearctic, Oriental, and New Guinean Pinophilina (Coleoptera: Staphylinidae: Paederinae)

    Volker Assing

    The Palaearctic, Oriental, and New Guinean fauna of the Pinophilina was previously represented by 78 valid species and one subspecies in seven genera, with 68 (87 %) of the species in the nominal genus Pinophilus Gravenhorst, 1802. This paederine subtribe had never been subject to a comprehensive revision. In order to assess...

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    The Palaearctic, Oriental, and New Guinean fauna of the Pinophilina was previously represented by 78 valid species and one subspecies in seven genera, with 68 (87 %) of the species in the nominal genus Pinophilus Gravenhorst, 1802. This paederine subtribe had never been subject to a comprehensive revision. In order to assess the taxonomy, systematics, and diversity of the Pinophilina in the study region, the accessible type material of previously named species, as well as numerous additional specimens, both previously identified and unidentified, from the major public and private collections were examined. In addition, relevant genera and species, in particular Pinophilus (type genus of Pinophilina) and Pinophilus latipes Gravenhorst, 1802 (type species of Pinophilus), from other zoogeographic regions were included in the study. The revision revealed that Pinophilus is absent from the Palaearctic and Oriental regions and probably confined to the New World. The subtribe is represented in the Palaearctic and Oriental regions, including New Guinea, by as many as 309 named species in 16 genera, and none of the previously described species remains in Pinophilus. Numerous species groups are newly established particularly in speciose genera. Except for two recently described species and one species whose type material is inaccessible, all the species are (re-)described and illustrated. | Eleven genera and 241 species are described for the first time: Mimopinophilus compar sp. nov. (Iran; Oman; Pakistan; India); M. hamatus sp. nov. (Indonesia: Sulawesi); M. triangularis sp. nov. (South India; Sri Lanka); M. uncinatus sp. nov. (Indonesia: Sumbawa); Neopinophilus abscisus sp. nov. (Malaysia: Selangor); N. crockermontis sp. nov. (Malaysia: Sabah); N. edentulus sp. nov. (South India); N. fulgens sp. nov. (Malaysia: Sarawak); N. glaber sp. nov. (Malaysia: Sarawak); N. kinabaluicus sp. nov. (Malaysia: Sabah); N. minodoronus sp. nov. (Philippines: Mindoro); N. muluanus sp. nov. (Malaysia: Sarawak); N. palawanus sp. nov. (Philippines: Palawan); N. rubellus sp. nov. (Indonesia: Sumatra); N. sandakanus sp. nov. (Malaysia: Sabah); N. selatanus sp. nov. (Indonesia: Sulawesi); N. tensicollis sp. nov. (Indonesia: Sulawesi); N. umbilicatus sp. nov. (Sri Lanka); Phinopilus acutior sp. nov. (China: Sichuan, Zhejiang); P. adclinis sp. nov. (China: Yunnan; Laos; Vietnam); P. adesus sp. nov. (Papua New Guinea); P. alishanus sp. nov. (Taiwan); P. ambangus sp. nov. (Indonesia: Sulawesi); P. ambulans sp. nov. (Philippines: Leyte); P. ampliatus sp. nov. (Papua New Guinea); P. amplicornis sp. nov. (North India); P. angustior sp. nov. (Indonesia: Kalimantan Selatan); P. arcticornis sp. nov. (Papua New Guinea); P. arquatus sp. nov. (Indonesia: Java); P. artior sp. nov. (China: Fujian, Zhejiang); P. auriculatus sp. nov. (North India); P. aviformis sp. nov. (Singapore); P. baculatus sp. nov. (Malaysia; Indonesia: Sumatra); P. biacifer sp. nov. (Sri Lanka); P. bilamellatus sp. nov. (China: Sichuan); P. boops sp. nov. (Malaysia: Sarawak); P. brevior sp. nov. (Malaysia: Sarawak); P. cacharicus sp. nov. (Northeast India); P. cacuminatus sp. nov. (Thailand; Laos; Cambodia); P. callicollis sp. nov. (Papua New Guinea, New Ireland, New Britain; Indonesia: Maluku Islands); P. candens sp. nov. (Malaysia; Indonesia); P. caninus sp. nov. (Thailand; Laos); P. celebesicus sp. nov. (Indonesia: Sulawesi); P. cernens sp. nov. (Malaysia: Sabah); P. chiangicus sp. nov. (Thailand); P. clavifer sp. nov. (Malaysia: Sabah); P. clinatus sp. nov. (New Guinea); P. coauditus sp. nov. (Thailand); P. collevatus sp. nov. (Western New Guinea; Papua New Guinea); P. confossus sp. nov. (Philippines: Palawan); P. cordiformis sp. nov. (South India); P. crenatus sp. nov. (Indonesia: Western New Guinea); P. cubans sp. nov. (Malaysia: Sabah); P. cultellatus sp. nov. (Brunei); P. curtatus sp. nov. (South India); P. digitalis sp. nov. (Cambodia); P. edolatus sp. nov. (Indonesia: Western New Guinea); P. eminens sp. nov. (Malaysia: Peninsular, Sabah, Sarawak); P. esuriens sp. nov. (Malaysia: Sarawak); P. excavatus sp. nov. (Cambodia); P. fibulatus sp. nov. (Thailand); P. figens sp. nov. (Indonesia: Java); P. firmicornis sp. nov. (Malaysia); P. fodens sp. nov. (Thailand); P. forticornis sp. nov. (Indonesia: Sumatra); P. garonus sp. nov. (India: Meghalaya); P. gradiens sp. nov. (Indonesia: North Maluku); P. grossicornis sp. nov. (Malaysia); P. hainanus sp. nov. (China: Hainan); P. hastatus sp. nov. (Vietnam); P. hebes sp. nov. (Malaysia: Sarawak; Brunei); P. inspicatus sp. nov. (Nepal; India; Myanmar; China; Thailand; Laos; Vietnam; Indonesia: Sumatra); P. lamellatus sp. nov. (Thailand); P. latibasalis sp. nov. (Sri Lanka); P. laticuspis sp. nov. (Indonesia: Western New Guinea); P. latior sp. nov. (Malaysia; Brunei); P. limans sp. nov. (Indonesia: Western New Guinea); P. linguatus sp. nov. (Brunei); P. linguiformis sp. nov. (Laos); P. loebli sp. nov. (Southwest India); P. longalatus sp. nov. (Indonesia: Sumatra); P. longastatus sp. nov. (Philippines: Luzon); P. longilobatus sp. nov. (Philippines: Leyte; Indonesia: Sulawesi); P. longipalpis sp. nov. (China: Yunnan); P. lunatus sp. nov. (Sri Lanka); P. monstrans sp. nov. (Nepal; Northeast India); P. peniculatus sp. nov. (Malaysia; Indonesia); P. perfodens sp. nov. (Thailand; Malaysia); P. petiolatus sp. nov. (Myanmar); P. reflexus sp. nov. (India; Myanmar; China; Thailand; Laos; Vietnam); P. robusticornis sp. nov. (Papua New Guinea); P. rossii sp. nov. (Thailand; Laos; Cambodia; Vietnam); P. rostratus sp. nov. (Southwest India); P. rotundatus sp. nov. (Indonesia: Western New Guinea); P. rubidipennis sp. nov. (Northwest India); P. schawalleri sp. nov. (Thailand; Laos); P. schillhammeri sp. nov. (Indonesia: Sumatra); P. shavrini sp. nov. (Philippines); P. solomonicus sp. nov. (Solomon Islands); P. spectans sp. nov. (Brunei); P. spiralis sp. nov. (Sri Lanka); P. substrictus sp. nov. (Solomon Islands; Papua New Guinea); P. subvexus sp. nov. (Laos); P. sulawesicus sp. nov. (Indonesia: Sulawesi); P. tamilus sp. nov. (South India); P. taurus sp. nov. (Malaysia: Sabah, Sarawak; Brunei); P. tensissimus sp. nov. (Nepal; India); P. torifer sp. nov. (Brunei); P. triacer sp. nov. (Papua New Guinea); P. tricuspis sp. nov. (Indonesia: Western New Guinea; Papua New Guinea); P. trifidus sp. nov. (Indonesia: Western New Guinea); P. trigonalis sp. nov. (India; Myanmar; Laos); P. trispinosus sp. nov. (Brunei); P. unguicus sp. nov. (Philippines: Palawan); P. unguifer sp. nov. (Indonesia: Sulawesi); P. unicuspis sp. nov. (New Guinea); P. validicornis sp. nov. (Malaysia: Sarawak), P. vates sp. nov. (Philippines: Leyte); P. vellicans sp. nov. (South India); P. vespertilio sp. nov. (Vietnam); P. vesperugo sp. nov. (Thailand; Cambodia); P. yunnanicus sp. nov. (China: Yunnan); Pinachenium gen. nov. (type species: Pinophilus longicornis Baudi di Selve, 1848); Pinobelus gen. nov. (type species: Pinophilus punctatissimus Sharp, 1889); Pinocavus gen. nov. (type species Pinocavus excisus sp. nov.; Saudi Arabia); P. volans sp. nov. (South India); Pinocharis acifera sp. nov. (Laos); P. biacuta sp. nov. (Myanmar); P. bifida sp. nov. (Thailand; Cambodia); P. burmana sp. nov. (Myanmar); P. clivis sp. nov. (Thailand; Laos; Cambodia; Vietnam); P. flagrans sp. nov. (South India; Sri Lanka); P. forans sp. nov. (India; Sri Lanka); P. furcillata sp. nov. (Laos); P. migra sp. nov. (Cambodia; Vietnam); P. proiecta sp. nov. (Thailand; Cambodia); P. unguica sp. nov. (Myanmar); P. velifera sp. nov. (Thailand; Laos; Cambodia; Vietnam); Pinocolis gen. nov. (type species: Pinophilus leucopus Kraatz, 1859); P. appendiculatus sp. nov. (Southwest India); P. asperifrons sp. nov. (Nepal; India: Assam); P. atrifemoralis sp. nov. (Philippines); P. bifilifer sp. nov. (South India); P. bisinuatus sp. nov. (China; Thailand; Laos; Vietam); P. commixtus sp. nov. (Myanmar); P. fuscitibialis sp. nov. (Hong Kong; China); P. gemellus sp. nov. (Myanmar); P. insectus sp. nov. (Indonesia: Kalimantan Selatan); P. mastigans sp. nov. (Thailand; Malaysia); P. piscator sp. nov. (South India); Pinocrenum gen. nov. (type species: Pinocrenum dentatum sp. nov.; Nepal); Pinonepalus aduncus sp. nov. (Malaysia: Perak); P. affimbriatus sp. nov. (Malaysia: Sabah); P. aquilus sp. nov. (Indonesia: Kalimantan Tengah); P. atricollis sp. nov. (Philippines: Luzon, Leyte); P. atricrus sp. nov. (Vietnam; Indonesia: Sumbawa, Lombok); P. auctus sp. nov. (Indonesia: Sulawesi); P. basoricus sp. nov. (Malaysia: Kelantan); P. bellipes sp. nov. (Malaysia: Sarawak; Indonesia: Kalimantan Tengah); P. bicarinatus sp. nov. (Malaysia: Sarawak); P. bicoloripes sp. nov. (Philippines: Mindanao); P. biconstrictus sp. nov. (Indonesia: Sumatra); P. biconvexus sp. nov. (Indonesia: Sumatra); P. brevilobatus sp. nov. (Indonesia: Sulawesi); P. claripes sp. nov. (Indonesia: Sumatra); P. coccinus sp. nov. (Indonesia: Sulawesi); P. contegens sp. nov. (Thailand); P. contractus sp. nov. (Malaysia); P. corculatus sp. nov. (Indonesia: Sulawesi); P. cordatus sp. nov. (Indonesia: Sulawesi); P. cybaeus sp. nov. (Philippines); Abstract 7 P. deambulans sp. nov. (Philippines: Mindanao); P. demorsus sp. nov. (Malaysia: Sabah); P. derectus sp. nov. (Malaysia: Sarawak); P. disparilis sp. nov. (Malaysia); P. distortus sp. nov. (Indonesia: Sumatra); P. districtus sp. nov. (Malaysia: Sabah); P. divexatus sp. nov. (Indonesia: Sumatra); P. elevatus sp. nov. (Sri Lanka); P. extensus sp. nov. (Indonesia: Sumatra); P. fissus sp. nov. (India: Meghalaya); P. fusus sp. nov. (China: Guangxi); P. gilvipes sp. nov. (Malaysia: Sabah); P. goanus sp. nov. (South India); P. hamulatus sp. nov. (Indonesia: Sumatra); P. hebetatus sp. nov. (Indonesia: Sulawesi); P. ingens sp. nov. (Indonesia: Sulawesi); P. iniquus sp. nov. (Thailand); P. latilobatus sp. nov. (Thailand); P. ligulatus sp. nov. (Philippines); P. limus sp. nov. (Malaysia); P. linguifer sp. nov. (Malaysia: Sarawak); P. luteipennis sp. nov. (Indonesia: Sumatra); P. luteipes sp. nov. (Philippines: Leyte); P. luzonicus sp. nov. (Philippines: Luzon); P. mancus sp. nov. (Malaysia); P. manasicus sp. nov. (India: Assam); P. migrus sp. nov. (Malaysia: Sabah); P. muluicus sp. nov. (Malaysia: Sarawak); P. muscicola sp. nov. (Malaysia: Sarawak); P. nigricrus sp. nov. (Thailand); P. ostentans sp. nov. (Philippines: Mindanao); P. palatus sp. nov. (Myanmar); P. parilis sp. nov. (Indonesia: Sumatra); P. pauli sp. nov. (Indonesia: Bali); P. pellucidus sp. nov. (Malaysia: Sabah, Sarawak); P. perlucens sp. nov. (Malaysia: Sarawak); P. polisicus sp. nov. (Philippines: Luzon); P. pravus sp. nov. (Indonesia: Sumatra); P. procerus sp. nov. (Philippines: Mindanao); P. pungens sp. nov. (Malaysia); P. retunsus sp. nov. (Indonesia: Sulawesi); P. sabahicus sp. nov. (Malaysia: Sabah); P. sagittans sp. nov. (Indonesia: Sumatra); P. sagittarius sp. nov. (Indonesia: Sumatra); P. siporanus sp. nov. (Indonesia: Metawei Islands); P. subtilior sp. nov. (Indonesia: Sulawesi); P. taleatus sp. nov. (Thailand); P. tortus sp. nov. (Malaysia: Selangor); P. translucidus sp. nov. (Malaysia: Sarawak); P. triquetrus sp. nov. (Laos; Vietnam); P. venetus sp. nov. (Malaysia: Sabah); Pinopellis gen. nov. (type species: Pinophilus borneensis Fauvel, 1895); Pinopellis apona sp. nov. (Philippines: Mindanao; P. aspera sp. nov. (Borneo: Brunei); P. biexcisa sp. nov. (Myanmar); P. breviata sp. nov. (Philippines: Mindanao, Leyte); P. burmana sp. nov. (Myanmar); P. clavulata sp. nov. (China: Jiangxi, ?Guizhou); P. laotica sp. nov. (Laos); P. longior sp. nov. (Malaysia: Pahang); P. mutabilis sp. nov. (Indonesia: Sulawesi); P. nepalensis sp. nov. (Nepal); P. russipes sp. nov. (Philippines: Luzon); P. spiculata sp. nov. (Philippines: Mindanao); P. tecta sp. nov. (Indonesia: Sulawesi); P. tensa sp. nov. (Indonesia: Sulawesi); P. tricarinata sp. nov. (Laos); P. truncata sp. nov. (Laos); P. turana sp. nov.(India: Meghalaya); P. volitans sp. nov. (Philippines: Mindanao); P. yunnanica sp. nov. (China: Yunnan); Pinoplanus gen. nov. (type species: Pinophilus aegyptius Erichson, 1840); Pinopressus gen. nov. (type species: Pinophilus complanatus Erichson, 1840); Pinoremus gen. nov. (type species: Pinoremus cuneatus sp. nov.; Papua New Guinea); Pinorima gen. nov. (type species: Pinorima spinosa sp. nov.; Thailand; China; Laos; Cambodia; Vietnam); Pinosenex gen. nov. (type species: Pinosenex falcatus sp. nov.; South India). In addition, one genus and four species from other zoogeographic regions are described: Pinoplanus spathulatus sp. nov. (Kenya; Namibia; Ethiopia); P. auctior sp. nov. (Tanzania); P. constrictus sp. nov. (South Africa; Zimbabwe); Novipinophilus gen. nov. (type species: Novipinophilus caledonicus sp. nov.; New Caledonia). Five names are revalidated: Pinocolis curtipennis (Bernhauer, 1935) (previously a synonym of Pinophilus femoratus Schubert, 1911); Pinocolis insignis (Sharp, 1874) (previously a synonym of Pinophilus javanus Erichson, 1840); Pinocolis pallipes (Kraatz, 1859) (previously a synonym of Pinophilus javanus); Pinoplanus densior (Bernhauer, 1935); Pinoplanus punctulatus (Fauvel, 1905). One genus-group and 17 species-group synonymies are proposed: Pinophilinus Eichelbaum, 1908 = Pinogalus Fagel, 1963, syn. nov.; Mimopinophilus formosae (Bernhauer, 1935) = M. coomani (Bernhauer, 1935), syn. nov.; Mimopinophilus picticornis (Kraatz, 1859) = M. laocoon (Fernando, 1959), syn. nov.; Mimopinophilus rufipennis (Sharp, 1874) = M. chinensis (Bernhauer, 1938), syn. nov.; Phinopilus sondaicus (Bernhauer, 1915) = P. proximatus (Cameron, 1936), syn. nov.; Pinachenium longicorne (Baudi di Selve, 1848) = P. depressum (Kraatz, 1859), syn. nov., = P. planum (Kraatz, 1859), syn. nov., = P. verticale (Fauvel, 1895), syn. nov., = P. wittmeri (Koch, 1934), syn. nov.; Pinobelus punctatissimus (Sharp, 1889) = P. parapunctatissimus (Li & Wang, 1993), syn. nov., = P. lini (Rougemont, 2017), syn. nov.; Pinocolis curtipennis (Bernhauer, 1935) = P. assamensis (Coiffait, 1982), syn. nov.; Pinocolis pallipes (Kraatz, 1859) = P. ferrugineus (Bernhauer, 1902), syn. nov.; Pinonepalus cassagnaui Coiffait, 1982 = P. nepalicus (Coiffait, 1982), syn. nov.; Pinonepalus rotundicollis (Kraatz, 1859) = P. orientalis (Cameron, 1920), syn. nov., = P. jacobsoni (Cameron, 1930), syn. nov., = P. perakanus (Cameron, 1950), syn. nov.; Pinopressus complanatus (Erichson, 1840) = P. ruficeps (Kraatz, 1859), syn. nov. In all, 104 new combinations are proposed, including 42 species distributed in the Afrotropical region: Mimopinophilus formosae (Bernhauer, 1935), comb. nov. (ex Pinophilus); Mimopinophilus rufipennis (Sharp, 1874), comb. nov. (ex Pinophilus); Mimopinophilus sanguinipennis (Bernhauer, 1935), comb. nov. (ex Pinophilus); Neopinophilus brachypterus (Kraatz, 1859), comb. nov. (ex Pinophilus); Neopinophilus funestus (Bernhauer, 1902), comb. nov. (ex Pinophilus); Phinopilus bomfordi (Eppelsheim, 1890), comb. nov. (ex Pinophilus); Phinopilus brevicornis (Cameron, 1952), comb. nov. (ex Pinophilus); Phinopilus brevis (Kraatz, 1859), comb. nov. (ex Pinophilus); Phinopilus ceylonicus (Cameron, 1931), comb. nov. (ex Pinophilus); Phinopilus drescheri (Cameron, 1936), comb. nov. (ex Pinophilus); Phinopilus falsus (Cameron, 1940), comb. nov. (ex Pinophilus); Phinopilus hindostanus (Cameron, 1945), comb. nov. (ex Pinophilus); Phinopilus lewisius (Sharp, 1874), comb. nov. (ex Pinophilus); Phinopilus papuanus (Cameron, 1937), comb. nov. (ex Pinophilus); Phinopilus peninsularis (Cameron, 1932), comb. nov. (ex Pinophilus); Phinopilus piceus (Fauvel, 1904), comb. nov. (ex Pinophilus); Phinopilus pilicollis (Motschulsky, 1858), comb. nov. (ex Pinophilus); Phinopilus preangeranus (Cameron, 1936), comb. nov. (ex Pinophilus); Phinopilus sachtlebeni (Bernhauer, 1935), comb. nov. (ex Pinophilus); Phinopilus sautteri (Bernhauer, 1935), comb. nov. (ex Pinophilus); Phinopilus senescens (Cameron, 1925), comb. nov. (ex Pinophilus); Phinopilus sondaicus (Bernhauer, 1915), comb. nov. (ex Pinophilus); Phinopilus uniformis (Cameron, 1928), comb. nov. (ex Pinophilus); Pinachenium longicorne (Baudi di Selve, 1949), comb. nov. (ex Pinophilus); Pinobelus punctatissimus (Sharp, 1889), comb. nov. (ex Pinophilus); Pinocavus abnormalis (Bernhauer, 1915), comb. nov. (ex Pinophilinus); Pinocavus africanus (Gestro, 1873), comb. nov. (ex Pinophilinus); Pinocavus auberti (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus australis (Gemminger & Harold, 1868), comb. nov. (ex Pinophilinus); Pinocavus gestroi (Gridelli, 1928), comb. nov. (ex Pinophilinus); Pinocavus gibbifrons (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus grandicollis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus itombwensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus kaboboensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus kahuziensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus lamottei (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus leleupi (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus luberoensis luberoensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus luberoensis dubius (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus minutus (Gridelli, 1928), comb. nov. (ex Pinophilinus); Pinocavus mwengensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus raffrayi (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus rugegensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus rugosus (Gridelli, 1928), comb. nov. (ex Pinophilinus); Pinocavus schatzmayri (Koch, 1934), comb. nov. (ex Pinophilinus); Pinocavus sjostedti (Eichelbaum, 1908), comb. nov. (ex Pinophilinus); 8 Abstract Pinocavus socotranus (Assing, 2012), comb. nov. (ex Pinophilinus); Pinocavus somalicus (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus spinosus (Assing, 2013), comb. nov. (ex Pinophilinus); Pinocavus strictipennis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocavus tshuapaensis (Fagel, 1963), comb. nov. (ex Pinophilinus); Pinocharis genalis (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinocharis rufoferruginea (Cameron, 1936), comb. nov. (ex Pinophilus); Pinocolis chapmani (Bernhauer, 1935), comb. nov. (ex Pinophilus); Pinocolis curtipennis (Bernhauer, 1935), comb. nov. (ex Pinophilus); Pinocolis femoratus (Schubert, 1911), comb. nov. (ex Pinophilus); Pinocolis insignis (Sharp, 1874), comb. nov. (ex Pinophilus); Pinocolis javanus (Erichson, 1840), comb. nov. (ex Pinophilus); Pinocolis leucopus (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinocolis malabarensis (Cameron, 1931), comb. nov. (ex Pinophilus); Pinocolis mixtus (Cameron, 1914), comb. nov. (ex Pinophilus); Pinocolis pallipes (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinocolis variipes (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinocolis vicinus (Cameron, 1931), comb. nov. (ex Pinophilus); Pinonepalus amicus (Bernhauer, 1915), comb. nov. (ex Pinophilus); Pinonepalus beccarii (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinonepalus bettotanus (Cameron, 1930), comb. nov. (ex Pinophilus); Pinonepalus brunneipennis (Cameron, 1932), comb. nov. (ex Pinophilus); Pinonepalus cameroni (Scheerpeltz, 1933), comb. nov. (ex Pinophilus); Pinonepalus eppelsheimi (Bernhauer, 1904), comb. nov. (ex Pinophilus); Pinonepalus melanocephalus (Motschulsky, 1858), comb. nov. (ex Pinophilus); Pinonepalus melanomerus (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinonepalus parvus (Bernhauer, 1914), comb. nov. (ex Pinophilus); Pinonepalus philippinus (Bernhauer, 1915), comb. nov. (ex Pinophilus); Pinonepalus rotundicollis (Kraatz, 1859), comb. nov. (ex Pinophilus); Pinonepalus selangorensis (Cameron, 1950), comb. nov. (ex Pinophilus); Pinonepalus separandus (Cameron, 1928), comb. nov. (ex Pinophilus); Pinonepalus thoracicus (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinopellis borneensis (Fauvel, 1895), comb. nov. (ex Pinophilus); Pinopellis coarcticollis (Cameron, 1941), comb. nov. (ex Pinophilus); Pinopellis nigripes (Cameron, 1914), comb. nov. (ex Pinophilus); Pinophilinus anosyanus (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus betschi (Jarrige, 1978), comb. nov. (ex Pinogalus); Pinophilinus brunneorufus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus cameroni (Fagel, 1963), comb. nov. (ex Pinogalus); Pinophilinus clementi (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus cordicollis (Jarrige, 1978), comb. nov. (ex Pinogalus); Pinophilinus daressalamensis (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus delkeskampi (Fagel, 1963), comb. nov. (ex Pinogalus); Pinophilinus fauveli (Schubert, 1902), comb. nov. (ex Pinogalus); Pinophilinus levasseuri (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus mantasoae (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus matumbianus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus micropterus (Bernhauer, 1937), comb. nov. (ex Pinogalus); Pinophilinus peyrierasi (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus rufus (Lecoq, 1986), comb. nov. (ex Pinogalus); Pinophilinus schuberti (Fauvel, 1905), comb. nov. (ex Pinogalus); Pinophilinus zambezianus (Fagel, 1963), comb. nov. (ex Pinogalus); Pinoplanus aegyptius (Erichson, 1840), comb. nov. (ex Pinophilus); Pinoplanus grandis (Reitter, 1906), comb. nov. (ex Pinophilus); Pinoplanus densior (Bernhauer, 1935), comb. nov.; Pinoplanus punctulatus (Fauvel, 1905), comb. nov.; Pinopressus complanatus (Erichson, 1840), comb. nov. (ex Pinophilus); Pinopressus indicus (Cameron, 1931), comb. nov. (ex Pinophilus). Mimopinophilus sanguinipennis (Bernhauer, 1935), previously a subspecies of M. formosae (Bernhauer, 1935) is regarded as a distinct species. A neotype is designated for Pinophilus siculus Kraatz, 1857. Lectotypes are designated for Pinophilus fossor Wollaston, 1868, P. zarudnyi Semenov, 1909, P. formosae Bernhauer, 1935, P. coomani Bernhauer, 1935, P. picticornis Kraatz, 1859, P. rufipennis Sharp, 1874, P. chinensis Bernhauer, 1938, P. sanguinipennis Bernhauer, 1935, P. punctatissimus Sharp, 1889, P. pallipes Kraatz, 1859, P. chapmani Bernhauer, 1935, P. variipes Fauvel, 1895, P. vicinus Cameron, 1931, P. malabarensis Cameron, 1931, P. curtipennis Bernhauer, 1935, P. mixtus Cameron, 1914, Pinophilus ruficeps Kraatz, 1859, P. indicus Cameron, 1931, P. depressus Kraatz, 1859, P. wittmeri Koch, 1934, P. brevicollis Erichson, 1840, P. bomfordi Eppelsheim, 1890, P. sachtlebeni Bernhauer, 1935, P. falsus Cameron, 1940, P. sautteri Bernhauer, 1935, P. ceylonicus Cameron, 1931, P. papuanus Cameron, 1937, P. uniformis Cameron, 1928, P. eppelsheimi Bernhauer, 1904, P. melanomerus Kraatz, 1859, P. rotundicollis Kraatz, 1859, P. amicus Bernhauer, 1915, P. philippinus Bernhauer, 1915, P. speculifrons Cameron, 1914, P. borneensis Fauvel, 1895, P. nigripes Cameron, 1914, P. coarcticollis Cameron, 1941, and P. genalis Fauvel, 1895, Numerous examined species are represented only by females and consequently not named. This and the general rarity of records suggest that a substantial number of species remains to be discovered and described in the future. A comprehensive catalogue and keys to the genera and species of Pinophilina of the Palaearctic and Oriental regions are provided. Aside from a derived aedeagal morphology, the genera represented in the study region share two evident synapomorphies: fused abdominal tergites IX and X and strongly dilated, compact protarsomeres I.IV (usually broader than combined length). The sole exception is Pinopressus, a genus characterised by an aedeagus of primitive condition, unfused tergites IX and X, and moderately dilated (not compact) protarsomeres I.IV. These observations suggest that Pinopressus forms a basal lineage within Pinophilina. At the generic level, there is currently no overlap between the fauna of the study region and that of the New World. On the other hand, many genera distributed in the Palaearctic and Oriental regions are present also in sub-Saharan Africa and/or Australia. Species diversity is highest in the warmer climates of the Oriental region, particularly in continental Southeast Asia and the Sunda Islands. With few exceptions (records with specified localities unknown), the distributions of all the species in the study region are mapped. Rather little is known regarding the natural history of Pinophilina. Most of the species were found at low to intermediate altitudes, few at high elevations (up to more than 3000 m). A substantial proportion of the species and specimens examined was collected primarily or exclusively at light, mostly close to lakes or rivers and streams. A number of species was exclusively extracted from litter and soil of forests, partly also from dead wood (bark, logs). The available ecological data and the general rarity of records suggest that the reproduction habitats are of a cryptic, partly probably subterranean nature.

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